Galvatron (Robots in Disguise)
Galvatron is Megatron reborn and much more powerful. In addition to his original modes he has gained four more- a dinosaur, a pteranodon, an elephant and a cruiser. What makes Galvatron so dangerous is the source of his powers- he drains power from others like an energy vampire, and the more power he steals, the stronger he becomes.
He first appeared on the scene to reclaim leadership of the Predacons from Scourge (who became sub-commander again), and prevent the Autobots from gaining control of Plasma (the head of Fortress Maximus). Galvatron was determined to gain control of Maximus through Scourge, who was linked with Plasma. Scourge finally succeeded in enslaving Fortress Maximus, and brought Galvatron and Sky-Byte to a demonstration, where Scourge revealed his true ambition and ordered Fortress Maximus to destroy both Galvatron and Sky-Byte. The Autobots showed up to turn the tide and attempt to defeat the Predacons. Galvatron stunned Scourge and told Sky-Byte to bring him and follow. The Autobots regained control of Fortress Maximus.
To prevent further treachery, Galvatron gathered Scourge and the other Decepticons and removed their sentience, turning them into little more than drones. He reinstated Sky-Byte as his sub-commander and ordered him to attack the Autobots' base and recapture Plasma. A fierce battle ensued with the Autobots seeming the victors until Galvatron entered the fray and brought the Predacon starship into it to engage Fortress Maximus in battle. Galvatron transformed into his hand mode and merged with his ship to fight Fortress Maximus. Maximus destroyed the ship, and seemingly slew Galvatron. But moments later, Galvatron emerged from the wreckage in his bat mode and started absorbing power from Fortress Maximus. When he was finished, all the Autobots together were no match for him, and were forced to retreat back into the Cybertron Net.
Galvatron sent the Decepticons after the Autobots and Sky-Byte followed. Galvatron used his power to flood the Cybertron Net with the magma deep beneath the surface of the Earth in an attempt to destroy all his enemies (if Sky-Byte perished as well, so be it). Ultra Magnus attempted to stop him, but he was no match for the Preadcon leader. Galvatron unleashed millions of tiny bat-shaped power siphons and sent them into the cities of Japan to draw power from humans, just as the Autobots saved themselves from his death trap and confronted him. But the siphons were finished drawing power from people and returned to Galvatron, where he absorbed the power and became even stronger and converted into the "Devilsaurer" (which he declared the ultimate combination of strength and speed).
He began to demonstrate his power, and it was all the Autobots could do to survive, as he was impervious to whatever they threw at him. He managed to imprison the Autobots in an energy net, and threatened to destroy them unless Omega Prime led him to what he considered a limitless source of power, the Earth's core. Omega Prime complied and led Galvatron to the core. Galvatron meant to show him his true power once he finished absorbing it and destroy Omega Prime. But Fortress Maximus managed to regain consciousness and channeled his power through the Cybertron Net straight to Prime. Omega Prime was able to unleash the power of the Matrix and created the gigantic Omega Fire Sword. Galvatron converted to robot mode and the two engaged in a final duel. Omega Prime defeated Galvatron, causing a huge explosion of all the power Galvatron had stolen. Galvatron was last seen banished in space along with his followers.
Japan
In Japan, Galvatron was known as Devil Gigatron, and his four new modes had specific names- the Devilsaurer, the Devilnodon, the Devilelephant and the Devilcruiser.